Played just about all of them in the OP.
The standouts are Smite and Awesomenauts.
Smite is far easier to control than the standard point-and-click games, and it has quite a few more options as far as battle movement is concerned.
Something that always bothered me in the rts-style MOBAs was the inability to run backwards while attacking an enemy. Because the only way to move is to click, and the only way to attack an enemy is to click, mouse movement is vital to success. If you play in a high resolution, you're at a disadvantage because you need to cover more screen space to achieve your actions.
Smite fixes this by binding movement to WASD and you use the mouse to attack, like a third-person shooter. This allows you to move in a direction while attacking in a different direction, which makes it play much faster and more easily than point-and-clicks.
Note: I'm sure you can bind other games to WASD, but I'm going by default settings.
The completely unique MOBA is Awesomenauts, though. Because it's 2d, jumping and vertical movement have a massive effect on the gameplay. In every other MOBA on the list, including Smite, jumping is merely cosmetic and doesn't really do anything. You play on a flat plane and move forward, backward, and side to side.
In Awesomenauts, you can move from the top level of the map to the bottom in a matter of seconds, which adds a lot to the strategy. You can move forward, backward, up and down, rather than side to side. This means you must be aware of your front, back, and attacks from below or above. It's much more 'twitch'-based, and it plays like oldschool arcade games.
If anybody is curious about which MOBA to start with, I'd definitely recommend Smite. It's free to play, and you can unlock characters via in-game currency, which is earned for doing well in matches. You can buy the characters, of course, but it's not vital to get every character.